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TCPIP bindings ~Hal Rekitexjip 22.Jan.04 04:44 AM a Web browser Domino Server 6.5Windows 2000
If I assume that Server 1 has two IP addresses (one for the internet/network and one for the cluster connection), you may find that the DNS lookups are trying to occur on the cluster network. This comes down to the port settings and order.
You can fix this by using notes.ini settings to bind SMTP to a specific port and therefore a specific IP address, where the DNS will function across correctly.
Try:
Ports=TCPINT,TCPEXT
TCPINT=TCP,0,15,0
TCPINT_TcpIpAddress=0,x.x.x.x:1352
TCPEXT=TCP,0,15,0
TCPEXT_TcpIpAddress=0,x.x.x.x:1352
SMTPNotesPort=TCPEXT